| 7,77911 % | Portolan Capital Management, L.L.C. | 3 633 622 | 670 225 | |
| 6,69522 % | Columbia Threadneedle Investments (UK) | 3 127 338 | 128 326 | |
| 5,66881 % | BlackRock Institutional Trust Company, N.A. | 2 647 901 | -36 107 | |
| 5,41415 % | The Vanguard Group, Inc. | 2 528 950 | 3 271 | |
| 4,47335 % | Kent Lake PR LLC | 2 089 500 | -211 500 | |
| 3,99109 % | Walt (David R) | 1 864 240 | 2 594 | |
| 2,45109 % | Invenomic Capital Management LP | 1 144 904 | 242 861 | |
| 2,41486 % | William Blair Investment Management, LLC | 1 127 981 | 21 918 | |
| 2,37538 % | Columbia Threadneedle Investments (US) | 1 109 539 | 393 313 | |
| 2,31992 % | Easterly Investment Partners LLC | 1 083 635 | -241 985 | |
| 2,05041 % | Blue Water Life Science Advisors, LP. | 957 747 | 585 572 | |
| 1,89912 % | Geode Capital Management, L.L.C. | 887 080 | 21 210 | |
| 1,72069 % | Dimensional Fund Advisors, L.P. | 803 735 | -37 598 | |
| 1,64376 % | Balyasny Asset Management LP | 767 801 | 723 295 | |
| 1,63745 % | State Street Investment Management (US) | 764 855 | 3 547 | |
| 1,47420 % | Trium Capital LLP | 688 597 | 102 172 | |
| 1,44726 % | Williams Jones Wealth Management LLC | 676 014 | 162 525 | |
| 1,35246 % | Morgan Stanley & Co. LLC | 631 732 | 180 440 | |
| 1,22376 % | Tikvah Management LLC | 571 619 | 0 | |
| 1,10467 % | Parkman Healthcare Partners LLC | 515 992 | 515 992 | |
| 0,95095 % | Marshall Wace LLP | 444 191 | 444 191 | |
| 0,90100 % | abrdn Inc. | 420 856 | 152 139 | |
| 0,82413 % | UBS Financial Services, Inc. | 384 951 | -730 027 | |
| 0,81168 % | Two Sigma Investments, LP | 379 137 | 229 674 | |
| 0,75859 % | SCS Capital Management LLC | 354 339 | 0 | |
| 0,69723 % | Schonfeld Strategic Advisors LLC | 325 677 | 325 677 | |
| 0,65464 % | Northern Trust Investments, Inc. | 305 784 | -2 754 | |
| 0,64621 % | Meister Paul M | 301 844 | 2 242 | |
| 0,64010 % | Charles Schwab Investment Management, Inc. | 298 989 | 6 184 | |
| 0,62034 % | State of Wisconsin Investment Board | 289 759 | 161 751 | |
| 0,61521 % | Toloue (Masoud) | 287 366 | -1 242 | |
| 0,60068 % | Citi Investment Research (US) | 280 578 | 254 082 | |
| 0,52045 % | J.P. Morgan Securities LLC | 243 101 | -27 609 | |
| 0,49583 % | Envestnet Asset Management, Inc. | 231 601 | -268 797 | |
| 0,32962 % | Goldman Sachs & Company, Inc. | 153 964 | -107 760 | |
| 0,32376 % | RBC Capital Markets Wealth Management | 151 230 | 2 261 | |
| 0,29964 % | Jane Street Capital, L.L.C. | 139 964 | 81 651 | |
| 0,29587 % | BlackRock Financial Management, Inc. | 138 200 | -78 230 | |
| 0,29573 % | Donnelly (William P) | 138 137 | 5 076 | |
| 0,29092 % | Parametric Portfolio Associates LLC | 135 888 | 112 032 | |
| 0,27112 % | Jump Financial, LLC | 126 641 | 21 184 | |
| 0,27110 % | Fidelity Management & Research Company LLC | 126 632 | 28 692 | |
| 0,26960 % | BlackRock Investment Management, LLC | 125 931 | 21 577 | |
| 0,26542 % | Empire Financial Management Company, LLC | 123 980 | 73 980 | |
| 0,25501 % | Wellington Management Company, LLP | 119 115 | 119 115 | |
| 0,25271 % | Cubist Systematic Strategies, LLC | 118 043 | -169 188 | |
| 0,24644 % | Wells Fargo Advisors | 115 113 | 91 688 | |
| 0,22094 % | Renaissance Technologies LLC | 103 200 | -99 721 | |
| 0,21409 % | ACATIS Investment Kapitalverwaltungsgesellschaft GmbH | 100 000 | 0 | |
| 0,21378 % | Goldman Sachs Asset Management, L.P. | 99 856 | 11 884 | |
| 0,18895 % | AQR Capital Management, LLC | 88 259 | -10 731 | |
| 0,18458 % | American Century Investment Management, Inc. | 86 215 | 7 085 | |
| 0,16756 % | Barclays Bank PLC | 78 266 | 24 818 | |
| 0,15313 % | Dimensional Fund Advisors, Ltd. | 71 529 | -4 725 | |
| 0,15213 % | Nuveen LLC | 71 058 | -342 523 | |
| 0,14751 % | BNY Mellon Asset Management | 68 902 | 3 080 | |
| 0,13916 % | Graypoint LLC | 65 000 | 0 | |
| 0,13912 % | Invesco Capital Management LLC | 64 984 | 21 167 | |
| 0,13771 % | Susquehanna International Group, LLP | 64 326 | 8 059 | |
| 0,13286 % | The Clarius Group, LLC | 62 060 | 0 | |
| 0,13231 % | BofA Global Research (US) | 61 801 | -75 594 | |
| 0,12980 % | Rhumbline Advisers Ltd. Partnership | 60 629 | 4 094 | |
| 0,12845 % | Mount Lucas Management LP | 60 000 | 16 163 | |
| 0,11994 % | XTX Markets LLC | 56 024 | 33 007 | |
| 0,11268 % | Hlavinka (Sarah E) | 52 635 | 2 615 | |
| 0,11155 % | Peddock Capital Advisors, LLC | 52 105 | 34 503 | |
| 0,10205 % | PDT Partners, LLC | 47 668 | 47 668 | |
| 0,09523 % | AllianceBernstein L.P. | 44 480 | 7 500 | |
| 0,09338 % | RBC Wealth Management, International | 43 616 | 21 840 | |
| 0,08984 % | Quantinno Capital Management LP | 41 964 | 31 664 | |
| 0,08526 % | Avantis Investors_NLE | 39 827 | 37 990 | |
| 0,08505 % | Mellon Investments Corporation | 39 726 | 1 802 | |
| 0,08478 % | T. Rowe Price International Ltd | 39 599 | 1 665 | |
| 0,07493 % | Credit Industriel et Commercial SA | 35 000 | 0 | |
| 0,07367 % | BNP Paribas Securities Corp. North America | 34 410 | -152 476 | |
| 0,06822 % | Quantbot Technologies, LP | 31 864 | 20 592 | |
| 0,06412 % | HSBC Global Asset Management (UK) Limited | 29 949 | -1 184 | |
| 0,06240 % | Nebula Research & Development LLC | 29 145 | -6 820 | |
| 0,05653 % | Flynn (Karen A) | 26 405 | 0 | |
| 0,05207 % | DWS Investments UK Limited | 24 321 | -6 394 | |
| 0,05102 % | MetLife Investment Management, LLC | 23 830 | -153 | |
| 0,05058 % | Wells Fargo Bank, N.A. | 23 624 | 2 669 | |
| 0,04559 % | Ironwood Investment Management, LLC | 21 297 | 21 297 | |
| 0,04389 % | Osaic Holdings, Inc. | 20 499 | 1 396 | |
| 0,04377 % | Mirae Asset Global Investments (USA) LLC | 20 444 | -936 | |
| 0,04332 % | Magovcevic-Liebisch (Ivana) | 20 235 | 7 435 | |
| 0,04282 % | Ersel Asset Management SGR S.p.A. | 20 000 | 0 | |
| 0,03859 % | T. Rowe Price Associates, Inc. | 18 025 | 5 300 | |
| 0,03768 % | Corebridge Financial Inc. | 17 602 | -262 | |
| 0,03739 % | Commonwealth Financial Network | 17 465 | -450 | |
| 0,03724 % | JPMorgan Private Bank (United States) | 17 397 | 405 | |
| 0,03617 % | INTECH Investment Management LLC | 16 893 | -6 585 | |
| 0,03615 % | Prelude Capital Management, LLC | 16 884 | -4 126 | |
| 0,03363 % | Tallon Kerry Patrick | 15 710 | 15 710 | |
| 0,03325 % | Metropolitan Life Insurance Co. (US) | 15 533 | 0 | |
| 0,03291 % | Squarepoint Capital LLP | 15 372 | -1 150 | |
| 0,03227 % | Sriram (Vandana) | 15 072 | -229 | |
| 0,03171 % | State of Wyoming | 14 814 | 7 599 | |
| 0,03105 % | Brevan Howard Capital Management LP | 14 505 | 14 505 | |
| 0,03094 % | Moors & Cabot Inc. | 14 452 | -18 001 | |